Overosaurus

Overosaurus Temporal range: Santonian ~86.3–83.6 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Overosaurus life restoration
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Dinosauria
Clade:	Saurischia
Clade:	†Sauropodomorpha
Clade:	†Sauropoda
Clade:	†Macronaria
Clade:	†Titanosauria
Clade:	†Lithostrotia
Genus:	†Overosaurus Coria et al., 2013
Species:	†O. paradasorum
Binomial name
†Overosaurus paradasorum Coria et al., 2013

Overosaurus (meaning "Overo lizard", after the Cerro Overo locality^[1]) is an extinct genus of sauropod dinosaurs, containing only a single species, Overosaurus paradasorum. This species lived approximately 86 to 84 million years ago during the latter part of the Cretaceous Period in what is now Patagonia (in southern Argentina). Overosaurus paradasorum was relatively small compared to other sauropods from Patagonia, like the saltasaurids and other aeolosaurines, estimated as approximately 10 m (33 ft).^[1] It was a ground-dwelling herbivore.

Discovery and naming

The only known specimen of Overosaurus paradasorum was recovered at the Cerro Overo locality in Patagonia, Argentina. The specimen was collected in 2002 by researchers from the Museo Carmen Funes, the Museo Argentino Urquiza, and the Natural History Museum of Los Angeles County, in terrestrial sediments deposited during the Campanian stage of the Late Cretaceous period, approximately 84 to 78 million years ago. This specimen is housed in the collection of the Museo Argentino Urquiza in Neuquén, Argentina. The specimen was found by Carlos Parada and family, who provided logistic support to museum staff^[2] as well as assistance during fieldwork.^[3] The Cerro Overo locality was originally reported as within the Anacleto Formation of the Neuquén Group,^[1] but is now attributed to the Bajo de la Carpa Formation.^[4]

Description

Overosaurus was described and named by Rodolfo A. Coria, Leonardo S. Filippi, Luis M. Chiappe, Rodolfo García and Andrea B. Arcucci in 2013. This species is known solely from the holotype MAU-Pv-CO-439 which consists of a fully articulated vertebral series from the 10th cervical to the 20th caudal vertebra, the ribs of the last three cervical vertebrae, six right dorsal ribs (articulated with their respective dorsal vertebrae 2,3,4,5,8,9), five left dorsal ribs (articulated with their respective dorsal vertebrae 2,3,4,5,8), the proximal portions of dorsal ribs of both sides of dorsal vertebrae 9 and 10, a complete right ilium and a fragmented left ilium. Specifically the vertebral series includes the last four cervical vertebrae, ten dorsal vertebrae, six sacral vertebrae, and twenty caudal vertebrae. Its sacrum consists of six fused sacral vertebrae.^[1]

Classification

Overosaurus was differentiated from other titanosaur taxa using a unique combination of characters present in the ilium and in the cervical, dorsal and caudal vertebrae. A phylogenetic analysis of Lithostrotia placed Overosaurus within the Aeolosaurini, as the sister taxon of a monophyletic group formed by Gondwanatitan faustoi, Pitekunsaurus macayai, Aeolosaurus rionegrinus, and Aeolosaurus maximus.

References

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Sauropodomorpha

Kingdom: Animalia Phylum: Chordata Class: Sauropsida Clade: Avemetatarsalia Clade: Dinosauria
Avemetatarsalia see Avemetatarsalia Sauropodomorpha see below↓

Sauropodomorpha
Sauropodomorpha Buriolestes Eoraptor Gigantoscelus Guaibasaurus Mbiresaurus Pampadromaeus Panphagia Saturnaliidae Buriolestes? Eoraptor? Chromogisaurus Nhandumirim Pampadromaeus? Panphagia? Saturnalia Bagualosauria Arcusaurus Asylosaurus Bagualosaurus Efraasia Nambalia Pantydraco Thecodontosaurus Unaysauridae Jaklapallisaurus Macrocollum Unaysaurus Plateosauria Plateosauravus Ruehleia Xixiposaurus Plateosauridae Euskelosaurus Gresslyosaurus Issi Pachysauriscus Plateosaurus Yimenosaurus Massopoda Gryponyx Gyposaurus Ignavusaurus Kholumolumo Musankwa Sarahsaurus Tuebingosaurus Riojasauridae Eucnemesaurus Riojasaurus Massospondylidae Adeopapposaurus Coloradisaurus Glacialisaurus Ignavusaurus? Leyesaurus Lufengosaurus Massospondylus Ngwevu Pradhania Sarahsaurus Xingxiulong? Sauropodiformes Chuxiongosaurus? Jingshanosaurus Qianlong Seitaad Xingxiulong? Yizhousaurus Yunnanosaurus Anchisauria Aardonyx Anchisaurus Camelotia Irisosaurus Lamplughsaura Leonerasaurus Melanorosaurus Meroktenos Mussaurus Pulanesaura Sefapanosaurus Sauropoda see below↓

Sauropoda
Sauropoda Amygdalodon Blikanasaurus Chinshakiangosaurus Gongxianosaurus Isanosaurus Leonerasaurus? Pulanesaura? Protognathosaurus Schleitheimia Tuebingosaurus? Lessemsauridae Antetonitrus Ingentia Ledumahadi Lessemsaurus Gravisauria Archaeodontosaurus? Kotasaurus Ohmdenosaurus Rhoetosaurus Sanpasaurus? Zizhongosaurus? Vulcanodontidae Tazoudasaurus Vulcanodon Eusauropoda Algoasaurus Asiatosaurus Bagualia Barapasaurus Bellusaurus Cetiosauriscus Dashanpusaurus Dystrophaeus Janenschia Jobaria Nebulasaurus Perijasaurus Shunosaurus Spinophorosaurus Tehuelchesaurus Volkheimeria Cetiosauridae Cetiosaurus Chebsaurus Ferganasaurus? Lapparentosaurus? Patagosaurus? Mamenchisauridae Analong Anhuilong Cetiosauriscus? Chuanjiesaurus Datousaurus? Eomamenchisaurus Huangshanlong Hudiesaurus Jingiella Klamelisaurus? Mamenchisaurus Omeisaurus Qijianglong Rhomaleopakhus Tienshanosaurus Tonganosaurus Wamweracaudia Xinjiangtitan Yuanmousaurus Zigongosaurus Turiasauria Amanzia Bellusaurus? Haestasaurus? Janenschia? Losillasaurus Mierasaurus Moabosaurus Narindasaurus Oplosaurus? Tehuelchesaurus? Tendaguria Turiasaurus Zby Neosauropoda Diplodocoidea (see below ↓ ) Macronaria (see below ↓ ) Dubious sauropods Bothriospondylus Cardiodon Dinodocus Gigantosaurus Neosodon Qinlingosaurus Ultrasaurus

Diplodocoidea
Diplodocoidea Haplocanthosaurus Diplodocimorpha Rebbachisauridae Agustinia Amazonsaurus Comahuesaurus Dzharatitanis? Histriasaurus Lavocatisaurus Maraapunisaurus? Sidersaura Xenoposeidon Zapalasaurus Khebbashia Limaysaurinae Cathartesaura Limaysaurus Nopcsaspondylus? Rayososaurus? Rebbachisaurinae Demandasaurus Itapeuasaurus Katepensaurus Nigersaurus Rebbachisaurus Tataouinea Flagellicaudata Dicraeosauridae Amargasaurus Amargatitanis Bajadasaurus Brachytrachelopan Dicraeosaurus Dyslocosaurus? Kaatedocus Lingwulong Pilmatueia Smitanosaurus Suuwassea Tharosaurus Diplodocidae Apatosaurinae Amphicoelias? Apatosaurus Atlantosaurus? Brontosaurus Diplodocinae Barosaurus Dinheirosaurus Diplodocus Galeamopus Kaatedocus? Leinkupal Supersaurus Tornieria

Macronaria
Macronaria Abrosaurus Aragosaurus Bashunosaurus? Daanosaurus Dashanpusaurus? Europasaurus Haestasaurus? Yuzhoulong Camarasauridae Camarasaurus Lourinhasaurus Titanosauriformes Astrodon Duriatitan Eucamerotus Fushanosaurus Fusuisaurus Ornithopsis Pelorosaurus Rugocaudia? Brachiosauridae Abydosaurus Atlasaurus Brachiosaurus Cedarosaurus Galvesaurus? Giraffatitan Lusotitan Sonorasaurus Soriatitan Venenosaurus Vouivria Somphospondyli Angolatitan Arkharavia Astrophocaudia Austrosaurus Brohisaurus Brontomerus Chubutisaurus Dongbeititan Europatitan Fukuititan Garumbatitan Jiangxititan? Jiutaisaurus Liaoningotitan Ligabuesaurus? Liubangosaurus Oceanotitan Padillasaurus Pukyongosaurus Ruixinia Sauroposeidon Sibirotitan Tastavinsaurus Triunfosaurus Euhelopodidae Australodocus? Chiayusaurus? Erketu Euhelopus Gannansaurus Gobititan Huabeisaurus Huanghetitan Jiangshanosaurus? Phuwiangosaurus Qiaowanlong Silutitan Tambatitanis Tangvayosaurus Yongjinglong Yunmenglong Diamantinasauria Australotitan Diamantinasaurus Sarmientosaurus Savannasaurus Wintonotitan? Titanosauria see below↓

Titanosauria
Titanosauria Abdarainurus Aegyptosaurus? Andesaurus Angolatitan? Arackar Argyrosaurus Atacamatitan Austroposeidon Baotianmansaurus Barrosasaurus Baurutitan? Bonatitan? Borealosaurus Brasilotitan Choconsaurus Daxiatitan Dongyangosaurus Dreadnoughtus? Elaltitan Epachthosaurus? Gandititan Hamititan Huabeisaurus Jiangshanosaurus? Kaijutitan Karongasaurus Laplatasaurus Ligabuesaurus? Magyarosaurus Malarguesaurus? Malawisaurus Mnyamawamtuka Narambuenatitan Ninjatitan Normanniasaurus Nullotitan Pellegrinisaurus? Petrobrasaurus Pitekunsaurus Rapetosaurus Rukwatitan Ruyangosaurus Tastavinsaurus? Tengrisaurus? Tiamat Traukutitan Trigonosaurus Sonidosaurus Uberabatitan Volgatitan Xianshanosaurus Diamantinasauria? Lirainosaurinae Ampelosaurus Atsinganosaurus Garrigatitan Lirainosaurus Lohuecotitan Mansourasaurus Paludititan Eutitanosauria Inawentu Menucocelsior Colossosauria Antarctosaurus Baalsaurus Bonitasaura Chucarosaurus Epachthosaurus? Jainosaurus Quetecsaurus Tengrisaurus? Vahiny Rinconsauria Adamantisaurus Maxakalisaurus? Muyelensaurus Overosaurus Panamericansaurus Rinconsaurus Aeolosaurini Aeolosaurus Arrudatitan Bravasaurus Caieiria Gondwanatitan Punatitan Shingopana Lognkosauria Argentinosaurus Drusilasaura Dzharatitanis? Futalognkosaurus Jiangxititan? Mendozasaurus Notocolossus Patagotitan Puertasaurus Saltasauroidea Bustingorrytitan Titanomachya Udelartitan Nemegtosauridae Nemegtosaurus Quaesitosaurus? Tapuiasaurus? Saltasauridae Igai Isisaurus? Qingxiusaurus Quaesitosaurus? Opisthocoelicaudiinae Alamosaurus Baurutitan? Dreadnoughtus? Opisthocoelicaudia Pellegrinisaurus? Zhuchengtitan? Saltasaurinae Abditosaurus Bonatitan? Ibirania Maxakalisaurus? Neuquensaurus Paralititan Rocasaurus Saltasaurus Yamanasaurus Dubious titanosaurs Bruhathkayosaurus? Campylodoniscus Clasmodosaurus Hypselosaurus Iuticosaurus Loricosaurus Macrurosaurus? Microcoelus Mongolosaurus? Titanosaurus

Topics in sauropodomorph research
Sauropod neck posture Sauropod hiatus List of sauropod species

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