Gueragama

Gueragama Temporal range: Aptian-Albian or Turonian-Campanian 125–100.5 Ma PreꞒ Ꞓ O S D C P T J K Pg N [1][2]
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Squamata
Suborder:	Iguania
Clade:	Acrodonta
Genus:	†Gueragama Simões et al., 2015[3]
Type species
†Gueragama sulamericana Simões et al., 2015[3]

Gueragama is an extinct genus of iguanian lizard from the Cretaceous of Brazil. It belongs to a group of iguanians called Acrodonta, whose living members include chameleons and agamids and are currently restricted to the Old World. Gueragama is the only acrodont known from South America, providing evidence that the group once ranged across much of Gondwana and only became restricted to the Old World after the supercontinent broke apart. The type species, Gueragama sulamericana, was named in 2015 on the basis of an isolated lower jaw from Goio-Erê Formation in the Bauru Basin, which was deposited in a desert environment. Although description shows age of Formation at Turonian to Campanian, later study proposes age around Aptian to Albian instead.^[1][2] Unlike modern acrodontans, whose teeth implant on the margins of the jaws, Gueragama has teeth that implant along the inner surface of the lower jaw, a feature common in most non-acrodontan lizards and characteristic of the second major group of iguanians, Pleurodonta. The non-acrodont dentition of Gueragama is evidence of its basal position within Acrodonta, and is shared with the taxon of Late Cretaceous acrodontan relatives, the family Priscagamidae.^[3] In 2022, it is considered that is unlikely to be acrodontan, more likely to be scincoids or teiids instead.^[4]

References

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Extinct squamate genera

• Kingdom: Animalia
• Phylum: Chordata
• Class: Reptilia
• Superorder: Lepidosauria

Iguania
Acrodonta Anqingosaurus Barbaturex Bharatagama Gueragama Xianglong Agamidae Indiagama Suratagama Pleurodonta / Iguanoidea Anolbanolis Suzanniwana Uquiasaurus Xihaina Corytophanidae Aciprion? Babibasiliscus Geiseltaliellus †Gobiguania Anchaurosaurus Ctenomastax Igua Isodontosaurus Polrussia Saichangurvel Temujinia Zapsosaurus Iguanidae Afairiguana Polychrotidae Armandisaurus Lapitiguana Pariguana Pristiguana Pumilia Phrynosomatidae Desertiguana Other extinct iguanians Brasiliguana Hoyalacerta? Huehuecuetzpalli? Swainiguanoides †Arretosauridae Arretosaurus Dornosaurus Ergiliinsaurus Khaichinguana †Priscagamidae Heterodontagama Mimeosaurus Phrynosomimus Pleurodontagama? Priscagama

†Polyglyphanodontia / Borioteiioidea

Chamopsiidae Chamops Gerontoseps Glyptogenys Harmodontosaurus Haptosphenus Leptochamops Meniscognathus Pelsochamops Socognathus Stypodontosaurus Trippenaculus Gilmoreteiidae / "macrocephalosaurines" Gilmoreteius Kuwajimalla Yechilacerta "Polyglyphanodontines" Bicuspidon Cherminsaurus Darchansaurus Distortodon Erdenetesaurus Polyglyphanodon Sineoamphisbaena? Other polyglyphanodontians Adamisaurus Funiusaurus Gobinatus Obamadon Olindalacerta? Paleochelco? Peneteius Tchingisaurus Tianyusaurus Tijubina? Tripennaculus

Stem Scincogekkonomorpha
Scincogekkonomorpha Ardeosaurus? Bavarisaurus? Eichstaettisaurus? Jucaraseps Liushusaurus? Pedrerasaurus Scandensia? Yabeinosaurus? Scleroglossa Gekkonomorpha Norellius Gekkota Cadurcogekko Cretaceogekko Gobekko Hoburogekko Yanatarogecko Stem Autarchoglossa (Evansauria) Parmeosaurus Tepexisaurus Tijubina? †Bainguidae? Bainguis? Eoxanta Globaura Myrmecodaptria

Scincomorpha

Cordyloidea Eocordyla Ornatocephalus Sakurasaurus Lacertoidea Cnodontosaurus Paracontogenys Purbicella †Polyglyphanodontia? Amphisbaenia Oligodontosaurus Todrasaurus Amphisbaenidae Listromycter Lophocranion Bipedidae Anniealexandria Blanidae Blanosaurus Cuvieribaena Louisamphisbaena †Chthonophidae Chthonophis Rhineuridae Archaerhineura Dyticonastis Hadrorhineura Hyporhina Jepsibaena Macrorhineura Ototriton Plesiorhineura Protorhineura Spathorhynchus †Polyodontobaenidae Polyodontobaena †Contogeniidae Contogenys Palaeoscincosaurus Utahgenys Lacertidae Amblyolacerta Camptognathosaurus? Dormaalisaurus Dracaenosaurus Escampcerta Gracilicerta Ligerosaurus Maioricalacerta Mediolacerta Plesiolacerta Pseudeumeces Pseudolacerta Quercycerta Teiidae Brevisaurus? Meyasaurus? Ptilotodon †Barbatteiidae Barbatteius Oardasaurus Xantusiidae Catactegenys Palepidophyma Palaeoxantusia Scincoidea Retinosaurus Scincoideus Slavoia Scincidae Axonoscincus Orthoscincus Pyrenasaurus? Other scincomorphs Apsgnathus Calanguban Chromatogenys Dakotaseps Kleskunsaurus Lonchisaurus Monocnemodon Pachygenys Saurillodon Saurillus Sciroseps Webbsaurus †Paramacellodidae Atokasaurus? Becklesius Mimobecklesisaurus Paramacellodus Parasaurillus? Pseudosaurillus? Sharovisaurus

Anguimorpha
Basal anguimorphs Chometokadmon Eolacerta Paramacellodus Paravaranus? Yabeinosaurus? †Mosasauroidea? †Aigialosauridae †Mosasauridae Neoanguimorpha Carusioidea Carusia Xenosauridae Entomophontes Exostinus Restes Anguidae Odaxosaurus Anguinae Headonhillia Helvetisaurus Ophisauriscus Ragesaurus Diploglossinae Apodosauriscus Eodiploglossus Gerrhonotinae Paragerrhonotus Parophisaurus †Glyptosaurinae Arpadosaurus Melanosaurus Gaultia Glyptosaurus Helodermoides Odaxosaurus Paraglyptosaurus Paraplacosauriops Peltosaurus Placosaurus Proglyptosaurus Proxestops Xestops Paleoanguimorpha Basal varanoids / platynotans Aiolosaurus Colpodontosaurus Dorsetisaurus Eosaniwa Palaeovaranus Parasaniwa Parviderma Proplatynotia Saniwides Telmasaurus Shinisauria? Bahndwivici Dalinghosaurus Merkurosaurus Provaranosaurus Goannasauria Basal Goannasaurians Asprosaurus Chianghsia Estesia Eurheloderma Gobiderma Lowesaurus Morohasaurus Palaeosaniwa Paraderma Primaderma Varanoidea Lanthanotidae Cherminotus Varanidae Ovoo Saniwa Archaeovaranus Other platynotans / varanoids Arcanosaurus? Cemeterius Dryadissector

Ophidia
Coniophis Diablophis Dinilysia? Eophis Lapparentophis Najash? Parviraptor Portugalophis Tetrapodophis See Serpentes (snakes)

Indeterminate squamates
Costasaurus Lamiasaura Oculudentavis? Schillerosaurus

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